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Family ARCHISARGIDAE
Archisargids are an extinct family of lower brachycerans known only from the Jurassic of Asia (China, Kazakhstan, and Mongolia) and one Jurassic species from New South Wales, Australia, the latter recently described by Oberprieler & Yeates (2012) and who also provided a list of most of the known species at that time. Archisargids were a fairly diverse family for flies of that time period, but only existed for a relatively short period of time geologically (less than 20 my; 145.4-164.7 mya). Previously treated as questionably included in the Eremochaetidae (Evenhuis, 1994b), Mostovski (1996a, 1996b, 1997) revised and raised it to family level transferring Archirhagio Rohdendorf, 1938, Mesophantasma Rohdendorf, 1962, Mesosolva Hong, 1983 and Prosolva Hong, 1983 to it. Previous to Mostovski's revision, the family was known only from one fossil species (Archisargus pulcher Rohdendorf, 1938). It has since then been the subject of many studies including a recent one by Zhang (2012b) who corrected the family identification of one of the early "tabanoid" pollinator genera described by Ren (1998) (Allomyia Ren) as an archisargid and not a tabanid. Zhang et al. (2011) proposed the family Origoasilidae for the new genus Origoasilus. This has since been shown by Zhang (2012c) to belong in the Archisargidae, where it is treated here.
In addition to the described taxa below, Ross & Jarzembowski (1996) and Coram & Jepson (2012) referred to a specimen from the Lower Cretaceous Purbeck formation in the UK as possibly belonging to this family. Currently, the Archisargidae are known from 61 species in 15 genera.
Ref.: Oberprieler & Yeates (2012, checklist of species); Zhang (2014, checklist of species).
Genus ARCHIRHAGIO Rohdendorf
ARCHIRHAGIO Rohdendorf, 1938: 35. Type species: Archirhagio obscurus Rohdendorf, 1938, by original designation.
gracilientus Wang, Shih, Ren & Wang, 2017: 232. PA: China (Middle Jurassic) [C].
obscurus Rohdendorf, 1938: 37. PA: Kazakhstan (Middle/Upper Jurassic) [C].
maximus Mostovski, 1997: 74 (75). PA: Kazakhstan (Middle/Upper Jurassic) [C].
striatus Zhang & Zhang, 2003: 74. PA: China (Middle Jurassic) [C].
varius Zhang, 2014: 5. PA: China (Middle Jurassic) [C].
zhangi Zhang, Yang & Ren in Zhang, Li, Yang & Ren, 2009: 62. PA: China (Middle Jurassic) [C].
mostovskii Zhang, 2014: 4. PA: China (Middle Jurassic) [C].
Genus ARCHISARGUS Rohdendorf
ARCHISARGUS Rohdendorf, 1938: 30. Type species: Archisargus pulcher Rohdendorf, 1938, by original designation.
pulcher Rohdendorf, 1938: 30. PA: Kazakhstan (Middle/Upper Jurassic) [C].
spurivenius Zhang, Yang, Ren & Shih, 2007a: 828. PA: China (Middle Jurassic) [C].
strigatus Zhang, Yang, Ren & Shih, 2007a: 830. PA: China (Middle Jurassic) [C].
Genus CALOSARGUS Mostovski
CALOSARGUS Mostovski, 1997: 74 (75). Type species: Calosargus tatianae Mostovski, 1997, by original designation.
Subgenus CALOSARGUS Mostovski
CALOSARGUS Mostovski, 1997: 74 (75) (as genus). Type species: Calosargus tatianae Mostovski, 1997, by original designation.
antiquus Zhang, Yang & Ren in Zhang, Yang, Ren & Shih, 2007b: 6. PA: China (Middle Jurassic) [C].
bellus Zhang, Yang & Ren in Zhang, Yang, Ren & Shih, 2007b: 3. PA: China (Middle Jurassic) [C].
daohugouensis Zhang, Yang & Re in Zhang, Yang, Ren & Shihn, 2007b: 9. PA: China (Middle Jurassic) [C].
hani Zhang, Yang & Ren in Zhang, Yang, Ren & Shih, 2007b: 15. PA: China (Middle Jurassic) [C].
niger Mostovski, 1997: 75 (76). PA: Kazakhstan (Middle/Upper Jurassic) [C].
talbragarensis Oberprieler & Yeates, 2012: 642. AU: Australia (Upper Jurassic)
tatianae Mostovski, 1997: 75 (76). PA: Kazakhstan (Middle/Upper Jurassic) [C].
tenuicellulatus Zhang, Yang & Ren in Zhang, Yang, Ren & Shih, 2007b: 10. PA: China (Middle Jurassic) [C].
validus Zhang, Yang & Ren in Zhang, Yang, Ren & Shih, 2007b: 13. PA: China (Middle Jurassic) [C].
Subgenus PTEROSARGUS Mostovski
PTEROSARGUS Mostovski, 1997: 75 (76). Type species: Calosargus (Pterosargus) thanasymus Mostovski, 1997, by original designation.
sinicus Zhang, 2010a: 312. PA: China (Middle Jurassic) [C].
thanasymus Mostovski, 1997: 75 (76). PA: Kazakhstan (Middle/Upper Jurassic) [C].
Genus DAOHUGOSARGUS Zhang
DAOHUGOSARGUS Zhang, 2012b: 66. Type species: Sharasargus eximius Zhang, Yang & Ren, 2008, by original designation.
eximius Zhang, Yang & Ren, 2008a: 270 (Sharasargus). PA: China (Middle Jurassic) [C].
Genus FLAGELLISARGUS Zhang
FLAGELLISARGUS Zhang, 2012c: 879. Type species: Flagellisargus sinicus Zhang, 2012, by original designation.
robustus Zhang, 2012c: 881. PA: China (Middle Jurassic) [C].
sinicus Zhang, 2012c: 879. PA: China (Middle Jurassic) [C].
venustus Zhang, 2012c: 880. PA: China (Middle Jurassic) [C].
Genus MESOPHANTASMA Rohdendorf
MESOPHANTASMA Rohdendorf, 1962: 319 (1991: 462). Type species: Mesophantasma tipuliforme Rohdendorf, 1962, by original designation.
tipuliforme Rohdendorf, 1962: 319 (1991: 462). PA: Kazakhstan (Middle/Upper Jurassic) [C].
Genus MESOSOLVA Hong
MESOSOLVA Hong, 1983: 133. Type species: Mesosolva parva Hong, 1983, by original designation.
PROSOLVA Hong, 1983: 135. Type species: Prosolva huabensis Hong, 1983, by original designation.
BREVISOLVA Zhang, Ren & Shih in Zhang, Yang, Ren & Shih, 2010a: 78. Type species: Brevisolva daohugouensis Zhang, Ren & Shih, 2010, by original designation.
angustocellulata Mostovski, 1996: 330. PA: Kazakhstan (Middle/Upper Jurassic) [C].
balyshevae Mostovski, 1996: 331. PA: Kazakhstan (Middle/Upper Jurassic) [C].
daohugouensis Zhang & Zhang, 2003: 73. PA: China (Middle Jurassic) [C].
dolosa Mostovski, 1996: 330. PA: Kazakhstan (Middle/Upper Jurassic) [C].
hennigi Mostovski, 1996: 330. PA: Kazakhstan (Middle/Upper Jurassic) [C].
huabeinsis Hong, 1983: 135 (Prosolva). PA: China (Middle Jurassic) [C].
imperfecta Mostovski, 1996: 331. PA: Kazakhstan (Middle/Upper Jurassic) [C].
jurassica Zhang, Yang & Shih in Zhang, Yang, Ren & Shih, 2010: 77. PA: China (Middle Jurassic) [C].
karataviensis Mostovski, 1996c: 120 (Prosolva). PA: Kazakhstan (Middle/Upper Jurassic) [C].
longivena Mostovski, 1996: 331. PA: Mongolia (Middle/Upper Jurassic) [C].
parva Hong, 1983: 133. PA: China (Middle Jurassic) [C].
rohdendorfi Mostovski, 1996: 329. PA: Kazakhstan (Middle/Upper Jurassic) [C].
sinensis Zhang, Yang & Ren in Zhang, Yang, Ren & Shih, 2010: 76. PA: China (Middle Jurassic) [C].
zhangae Zhang, 2012: 69 (new replacement name for daohugouensis Zhang, Ren & Shih in Zhang, Yang, Ren & Shih, 2010a).
daohugouensis Zhang, Ren & Shih in Zhang, Yang, Ren & Shih, 2010a: 79 (Brevisolva). PA: China (Middle Jurassic) [C]. [Preoccupied Zhang & Zheng, 2003; no new name proposed.]
Genus NOVISARGUS Zhang
NOVISARGUS Zhang, 2014: 9. Type species: Novisargus rarus Zhang, 2014, by original designation.
rarus Zhang, 2014: 9. PA: China (Middle Jurassic) [C].
Genus ORIGOASLIUS Zhang, Yang & Ren
ORIGOASLIUS Zhang, Yang & Ren, 2011: 994. Type species: Origoasilus pingquanensis Zhang, Yang & Ren, 2011, by original designation.
pingquanensis Zhang, Yang & Ren, 2011: 994. PA: China (Middle Jurassic) [C].
Genus OVISARGUS Mostovski
OVISARGUS Mostovski, 1996c: 121. Type species: Ovisargus gracilis Mostovski, 1996, by original designation.
Subgenus HELEMPIS Ren
HELEMPIS Ren, 1998: 80 (as genus). Type species: Helempis yixianensis Ren, 1998, by original designation.
eucalla Ren, 1998: 80 (Helempis). PA: China (Upper Jurassic) [C].
yixianensis Ren, 1998; 80 (Helempis). PA: China (Upper Jurassic) [C].
Subgenus OVISARGUS Mostovski
gracilis Mostovski, 1996c: 121. PA: Kazakhstan (Middle/Upper Jurassic) [C].
singulus Zhang, 2014: 11. PA: China (Middle Jurassic) [C].
Genus PARVISARGUS Mostovski
PARVISARGUS Mostovski, 1996c: 122. Type species: Parvisargus malus Mostovski, 1996, by original designation.
malus Mostovski, 1996c: 123. PA: Kazakhstan (Middle/Upper Jurassic) [C].
peior Mostovski, 1996c: 123. PA: Kazakhstan (Middle/Upper Jurassic) [C].
Genus SHARASARGUS Mostovski
SHARASARGUS Mostovski, 1996c: 124. Type species: Sharasargus ruptus Mostovski, 1996, by original designation.
PAUROMYIA Ren, 1998: 72. Type species: Pauromyia oresbia Ren, 1998, by original designation.
fortis Zhang, Yang & Ren, 2008b: 271. PA: China (Middle Jurassic) [C].
maculus Zhang, 2014: 12. PA: China (Middle Jurassic) [C].
oresbius Ren, 1998; 71 (Pauromyia). PA: China (Upper Jurassic) [C].
ruptus Mostovski, 1996c: 124. PA: Mongolia (Upper Jurassic) [C].
spiniger Mostovski, 1996c: 122. PA: Kazakhstan (Middle/Upper Jurassic) [C].
Genus SINALLOMYIA Zhang
ALLOMYIA Ren, 1998: 68. Type species: Allomyia ruderalis Ren, 1998, by original designation. [Preoccupied by Banks, 1916.]
SINALLOMYIA Zhang, 2012b: 66 (new replacement name for Allomyia Ren, 1998). Type species: Allomyia ruderalis Ren, 1998, automatic.
ruderalis Ren, 1998; 68 (Allomyia). PA: China (Upper Jurassic) [C].
Genus TABANISARGUS Zhang
TABANISARGUS Zhang, 2014: 15. Type species: Tabanisargus daohugouensis Zhang, 2014, by original designation.
daohugouensis Zhang, 2014: 15. PA: China (Middle Jurassic) [C].
Genus URANORHAGIO Zhang, Yang & Ren
URANORHAGIO Zhang, Yang & Ren, 2010b: 564. Type species: Uranorhagio daohugouensis Zhang, Yang & Ren, 2010, by original designation.
STENORHAGIO Zhang, Yang & Ren, 2010b: 566. Type species: Stenorhagio deviatus Zhang, Yang & Ren, 2010, by original designation.
MOSTOVSKISARGUS Zhang, 2010a: 310. Type species: Mostovskisargus portentosus Zhang, 2010, by original designation.
asymmetricus Zhang, Yang & Ren, 2010b: 568 (Stenorhagio). PA: China (Middle Jurassic) [C].
conjugovenius Zhang, Yang & Ren, 2010b: 569 (Stenorhagio). PA: China (Middle Jurassic [C].
daohugouensis Zhang, Yang & Ren, 2010b: 565. PA: China (Middle Jurassic) [C].
deviatus Zhang, Yang & Ren, 2010b: 567 (Stenorhagio). PA: China (Middle Jurassic) [C].
grimaldi Zhang, Yang & Ren, 2010b: 567 (Stenorhagio). PA: China (Middle Jurassic) [C].
portentosus Zhang, 2010a: 310 (Mostovskisargus). PA: China (Middle Jurassic) [C].
signatus Zhang, 2010a: 311 (Mostovskisargus). PA: China (Middle Jurassic) [C].